For me, you should really think of the season and the climat your want to use, this would be huge in term of story telling. Winter -> forest is dead, they are some kind of surviror, Spring -> reborn of the forest, let's kick out these men, Summer -> kind of a equivalent fight, Fall -> the forest is dying.
And I might say I love the fall for the story, so I would go over your last version which has a really good vibe for this